VivoSim Labs, INC. — Director's Dealing 2018

May 29, 2018

SEC Form 4 — Statement of Changes in Beneficial Ownership

Issuer: ORGANOVO HOLDINGS, INC. (ONVO)
CIK: 0001497253
Period of Report: 2018-05-24

Reporting Person: CROUCH TAYLOR (Director, CEO & President)

Non-Derivative Transactions

Date	Security	Code	Shares	Price	A/D	Holdings After	Ownership
2018-05-24	Common Stock	A	148760	$0.00	Acquired	148760	Direct

Derivative Transactions

Date	Security	Exercise Price	Code	Shares	A/D	Expiration	Underlying	Ownership
2018-05-24	Stock Option (Right to Buy)	$1.84	A	600000	Acquired	2028-05-24	Common Stock (600000)	Direct

Footnotes

F1: Annual long-term equity award approved by the Compensation Committee of the Issuer's Board of Directors.

F2: The shares represent a grant of Restricted Stock Units ("RSUs"). The RSUs represent a right to receive shares of the Issuer's common stock, at no additional conversion or exercise price. The RSUs vest and settle over 16 equal quarterly installments measured from May 15, 2018.

F3: This number does not include 2,688,212 shares of common stock underlying outstanding options held by Mr. Crouch; post-transaction Mr. Crouch continues to beneficially own or have a right to acquire 2,836,972 shares of common stock.

F4: The option shares vest and become exercisable over 16 equal quarterly installments measured from May 15, 2018.