VivoSim Labs, INC. — Director's Dealing 2017

May 18, 2017

SEC Form 4 — Statement of Changes in Beneficial Ownership

Issuer: ORGANOVO HOLDINGS, INC. (ONVO)
CIK: 0001497253
Period of Report: 2017-05-16

Reporting Person: Presnell Sharon Collins (Chief Scientific Officer)

Non-Derivative Transactions

Date	Security	Code	Shares	Price	A/D	Holdings After	Ownership
2017-05-16	Common Stock	F	1167	$2.8621	Disposed	99188	Direct

Footnotes

F1: Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in July 2016, pursuant to the Reporting Person's agreement with the Issuer.

F2: This number does not include 1,282,192 shares of common stock underlying outstanding options held by Dr. Presnell; post-transaction Dr. Presnell continues to beneficially own or have a right to acquire 1,381,380 shares of common stock.